You may notice that you obtain Skull Runes after you level up and upon completing battlegrounds. The further you level up the more Skull Runes you will be given as you progress.
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In order to use these Skull Runes, you must first build The Obelisk of Power to access the Rune Tree. The Rune Tree allows you to place Runes in order to get certain upgrades for your units. When you first start you must use two Skull Runes to unlock Unit Set 1.
As you progress down the tree, upgrades will successively take more time to complete. In addition, you will have to unlock the previous upgrade before going further down the tree.
For example, before you can unlock Dwarf Speed you must first get Pikeman Speed to Level 8. The same goes for Pikeman Speed, with its prerequisite being Archer Speed Level 8.
Note: You can be upgrading at the Obelisk of Power, while completing other tasks at the same time.
The last thing about Skull Runes is VERY IMPORTANT. In the top right corner in the Obelisk of Power you will see an "Edit" button. This will allow you to remove your Skull Runes and redistribute them for free. However, this can only be done ONCE. After the first time you click Edit it will cost you more and more Sapphire each time you want to redistribute your Skull Runes. Make sure you plan out your Skull Rune allocation carefully.
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